P. Abbate, F. Andrade, and J. Culot, The effects of radiation and nitrogen on number of grains in wheat, The Journal of Agricultural Science, vol.23, issue.03, pp.351-360, 1995.
DOI : 10.1016/0378-4290(89)90071-3

M. Rousseaux, A. Hall, and R. Sanchez, Far-red enrichment and photosynthetically active radiation level influence leaf senescence in field-grown sunflower, Physiologia Plantarum, vol.80, issue.2, pp.217-224, 1996.
DOI : 10.1016/0378-4290(88)90008-1

V. Sadras, A. Hall, and D. Connor, Light-associated nitrogen distribution profile in flowering canopies of sunflower (Helianthus annuus L.) altered during grain growth, Oecologia, vol.80, issue.4, pp.488-494, 1993.
DOI : 10.1007/BF00317432

T. Shiraiwa and T. Sinclair, Distribution of Nitrogen among Leaves in Soybean Canopies, Crop Science, vol.33, issue.4, pp.804-808, 1993.
DOI : 10.2135/cropsci1993.0011183X003300040035x

B. Singh and C. Jenner, A modified method for the determination of cell number in wheat endosperm, Plant Science Letters, vol.26, issue.2-3, pp.273-278, 1982.
DOI : 10.1016/0304-4211(82)90101-8

T. Udagawa, Z. Uchijima, T. Horie, and K. Kobayashi, Studies of Energy and Gas Exchange within Crop Canopies (3) : Canopy structure of corn plants, Japanese journal of crop science, vol.37, issue.4, pp.589-597, 1968.
DOI : 10.1626/jcs.37.589

Z. Wang, A. Wei, and D. Zheng, Photosynthetic characteristics of non-leaf organs of winter wheat cultivars differing in ear type and their relationship with grain mass per ear, Photosynthetica, vol.39, issue.2, pp.239-244, 2001.
DOI : 10.1023/A:1013743523029

L. Weaver and R. Amasino, Senescence Is Induced in Individually Darkened Arabidopsis Leaves, but Inhibited in Whole Darkened Plants, PLANT PHYSIOLOGY, vol.127, issue.3, pp.876-886, 2001.
DOI : 10.1104/pp.010312

. Literature, N. Anten, T. Hirose, Y. Onoda, T. Kinugasa et al., Elevated CO 2 and nitrogen availability have interactive effects on canopy carbon gain in rice, New Phytol, vol.161, pp.459-471, 2004.

N. Anten, F. Schieving, and M. Werger, Patterns of light and nitrogen distribution in relation to whole canopy carbon gain in C3 and C4 mono- and dicotyledonous species, Oecologia, vol.102, issue.4, pp.504-513, 1995.
DOI : 10.1007/BF00329431

J. Araus and L. Tapia, Photosynthetic Gas Exchange Characteristics of Wheat Flag Leaf Blades and Sheaths during Grain Filling: The Case of a Spring Crop Grown under Mediterranean Climate Conditions, PLANT PHYSIOLOGY, vol.85, issue.3, pp.667-673, 1987.
DOI : 10.1104/pp.85.3.667

R. Austin, Yield of Wheat in the United Kingdom, Crop Science, vol.39, issue.6, pp.1604-1610, 1999.
DOI : 10.2135/cropsci1999.3961604x

A. Barneix and M. Guitman, Leaf Regulation of the Nitrogen Concentration in the Grain of Wheat Plants, Journal of Experimental Botany, vol.44, issue.10, pp.1607-1612, 1993.
DOI : 10.1093/jxb/44.10.1607

J. Bertheloot, B. Andrieu, C. Fournier, and P. Martre, A process-based model to simulate nitrogen distribution within wheat (Triticum aestivum L.) during grain filling, Funct Plant Biol, 2008.

P. Bindraban, Impact of canopy nitrogen profile in wheat on growth, Field Crops Research, vol.63, issue.1, pp.63-77, 1999.
DOI : 10.1016/S0378-4290(99)00030-1

A. Boonman, E. Prinsen, F. Gilmer, U. Schurr, A. Peeters et al., Cytokinin Import Rate as a Signal for Photosynthetic Acclimation to Canopy Light Gradients, PLANT PHYSIOLOGY, vol.143, issue.4, pp.1841-1852, 2007.
DOI : 10.1104/pp.106.094631

A. Borrell, G. Hammer, and E. Van-oosterom, Stay-green: A consequence of the balance between supply and demand for nitrogen during grain filling?, Annals of Applied Biology, vol.80, issue.1, pp.91-95, 2001.
DOI : 10.1093/jexbot/51.suppl_1.329

D. Calderini, M. Dreccer, and G. Slafer, Consequences of breeding on biomass, radiation interception and radiation-use efficiency in wheat, Field Crops Research, vol.52, issue.3, pp.271-281, 1997.
DOI : 10.1016/S0378-4290(96)03465-X

J. Chen, J. Reynolds, P. Harley, and J. Tenhunen, Coordination theory of leaf nitrogen distribution in a canopy, Oecologia, vol.39, issue.1, pp.63-69, 1993.
DOI : 10.1007/BF00321192

D. Connor, V. Sadras, and A. Hall, Canopy nitrogen distribution and the photosynthetic performance of sunflower crops during grain filling ? a quantitative analysis, Oecologia, vol.44, issue.3, pp.274-281, 1995.
DOI : 10.1007/BF00328812

I. Donnison, A. Gay, H. Thomas, K. Edwards, D. Edwards et al., ) by transposon insertional mutagenesis in a protease gene, New Phytologist, vol.43, issue.3, pp.481-494, 2007.
DOI : 10.1111/j.1469-8137.2006.01928.x

M. Dreccer, G. Slafer, and R. Rabbinge, Optimization of Vertical Distribution of Canopy Nitrogen, Journal of Crop Production, vol.57, issue.1, pp.47-77, 1998.
DOI : 10.1016/0378-4290(93)90144-C

M. Dreccer, M. Van-oijen, A. Schapendonk, C. Pot, and R. Rabbinge, Dynamics of Vertical Leaf Nitrogen Distribution in a Vegetative Wheat Canopy. Impact on Canopy Photosynthesis, Annals of Botany, vol.86, issue.4, pp.821-831, 2000.
DOI : 10.1006/anbo.2000.1244

J. Drouet and R. Bonhomme, Do Variations in Local Leaf Irradiance Explain Changes to Leaf Nitrogen within Row Maize Canopies?, Annals of Botany, vol.84, issue.1, pp.61-69, 1999.
DOI : 10.1006/anbo.1999.0890

J. Drouet and R. Bonhomme, Effect of 3D Nitrogen, Dry Mass per Area and Local Irradiance on Canopy Photosynthesis Within Leaves of Contrasted Heterogeneous Maize Crops, Annals of Botany, vol.93, issue.6, pp.699-710, 2004.
DOI : 10.1093/aob/mch099

J. Evans, Partitioning of Nitrogen Between and Within Leaves Grown Under Different Irradiances, Australian Journal of Plant Physiology, vol.16, issue.6, pp.533-548, 1989.
DOI : 10.1071/PP9890533

U. Feller and A. Fischer, Nitrogen Metabolism in Senescing Leaves, Critical Reviews in Plant Sciences, vol.22, issue.3, pp.241-273, 1994.
DOI : 10.1093/jxb/38.7.1174

U. Feller, T. Soong, and R. Hageman, Leaf Proteolytic Activities and Senescence during Grain Development of Field-grown Corn (Zea mays L.), PLANT PHYSIOLOGY, vol.59, issue.2, 1977.
DOI : 10.1104/pp.59.2.290

C. Field, Allocating leaf nitrogen for the maximization of carbon gain: Leaf age as a control on the allocation program, Oecologia, vol.38, issue.2-3, pp.341-347, 1983.
DOI : 10.1007/BF00379710

A. Fischer, Nutrient Remobilization During Leaf Senescence, Annual Plant Reviews, vol.26, pp.87-107, 2007.
DOI : 10.1002/9780470988855.ch5

E. Frak, L. Roux, X. Millard, P. Adam, B. Dreyer et al., Spatial distribution of leaf nitrogen and photosynthetic capacity within the foliage of individual trees: disentangling the effects of local light quality, leaf irradiance, and transpiration, Journal of Experimental Botany, vol.53, issue.378, pp.2207-2216, 2002.
DOI : 10.1093/jxb/erf065
URL : https://hal.archives-ouvertes.fr/hal-00964646

K. Hikosaka, I. Terashima, and S. Katoh, Effects of leaf age, nitrogen nutrition and photon flux density on the distribution of nitrogen among leaves of a vine (Ipomoea tricolor Cav.) grown horizontally to avoid mutual shading of leaves, Oecologia, vol.97, issue.4, pp.451-457, 1994.
DOI : 10.1007/BF00325881

B. Hirel, B. Andrieu, M. Valadier, R. S. Quillere, I. Chelle et al., Physiology of maize II: Identification of physiological markers representative of the nitrogen status of maize (Zea mays) leaves during grain filling, Physiologia Plantarum, vol.41, issue.2, pp.178-188, 2005.
DOI : 10.1093/jexbot/53.370.917

T. Hirose and M. Werger, Maximizing daily canopy photosynthesis with respect to the leaf nitrogen allocation pattern in the canopy, Oecologia, vol.73, issue.4, pp.520-526, 1987.
DOI : 10.1007/BF00378977

L. Irving and D. Robinson, A dynamic model of Rubisco turnover in cereal leaves, New Phytologist, vol.162, issue.3, pp.493-504, 2006.
DOI : 10.1111/j.1469-8137.2005.01584.x

M. Kade, A. Barneix, S. Olmos, and J. Dubcovsky, Nitrogen uptake and remobilization in tetraploid 'Langdon' durum wheat and a recombinant substitution line with the high grain protein gene Gpc-B1, Plant Breeding, vol.36, issue.4, pp.343-349, 2005.
DOI : 10.1016/S1161-0301(01)00146-0

Y. Kato, S. Murakami, Y. Yamamoto, H. Chatani, Y. Kondo et al., The DNA-binding protease, CND41, and the degradation of ribulose-1,5-bisphosphate carboxylase/oxygenase in senescent leaves of tobacco, Planta, vol.338, issue.1, pp.97-104, 2004.
DOI : 10.1007/s00425-004-1328-0

H. Kim, M. Lieffering, S. Miura, K. Kobayashi, and M. Okada, Growth and nitrogen uptake of CO2-enriched rice under field conditions, New Phytologist, vol.89, issue.2, pp.223-229, 2001.
DOI : 10.1023/A:1004786414113

K. Kobayashi and M. Salam, Comparing Simulated and Measured Values Using Mean Squared Deviation and its Components, Agronomy Journal, vol.92, issue.2, pp.345-352, 2000.
DOI : 10.1007/s100870050043

O. Kull, Acclimation of photosynthesis in canopies: models and limitations, Oecologia, vol.133, issue.3, pp.267-279, 2002.
DOI : 10.1007/s00442-002-1042-1

O. Kull and B. Kruijt, Acclimation of photosynthesis to light: a mechanistic approach, Functional Ecology, vol.13, issue.1, pp.24-36, 1999.
DOI : 10.1146/annurev.arplant.40.1.119

A. Lang, Application of some of Cauchy's theorems to estimation of surface areas of leaves, needles and branches of plants, and light transmittance, Agricultural and Forest Meteorology, vol.55, issue.3-4, pp.191-212, 1991.
DOI : 10.1016/0168-1923(91)90062-U

M. Lö-tscher, K. Stroh, and H. Schnyder, Vertical Leaf Nitrogen Distribution in Relation to Nitrogen Status in Grassland Plants, Annals of Botany, vol.92, issue.5, pp.679-688, 2003.
DOI : 10.1093/aob/mcg188

A. Makino, T. Mae, and K. Ohiro, Relation between nitrogen and ribulose- 1,5-bisphosphate carboxylase in rice leaves from emergence through senescence, Plant Cell Physiol, vol.25, pp.429-437, 1984.

P. Martre, J. Porter, P. Jamieson, and T. , Modeling Grain Nitrogen Accumulation and Protein Composition to Understand the Sink/Source Regulations of Nitrogen Remobilization for Wheat, PLANT PHYSIOLOGY, vol.133, issue.4, pp.1959-1967, 2003.
DOI : 10.1104/pp.103.030585

P. Martre, M. Semenov, and P. Jamieson, Simulation Analysis of Physiological Traits to Improve Yield, Nitrogen Use Efficiency and Grain Protein Concentration in Wheat, Scale and Complexity in Plant Systems Research, Gene-Plant-Crop Relations, pp.181-201, 2007.
DOI : 10.1007/1-4020-5906-X_15

C. Masclaux-daubresse, E. Carrayol, and M. Valadier, The two nitrogen mobilisation- and senescence-associated GS1 and GDH genes are controlled by C and N metabolites, Planta, vol.46, issue.4, pp.580-588, 2005.
DOI : 10.1007/s00425-004-1468-2

M. Monsi and T. Saeki, On the Factor Light in Plant Communities and its Importance for Matter Production, Annals of Botany, vol.95, issue.3, pp.549-567, 2005.
DOI : 10.1093/aob/mci052

H. Mooney, S. Gulmon, G. Jain, and . Johnson, Environmental and Evolutionary Constraints on the Photosynthetic Characteristics of Higher Plants, Topics in Plant Population Biology, pp.316-337, 1979.
DOI : 10.1007/978-1-349-04627-0_14

P. Oscarson, T. Lundborg, M. Larsson, and C. Larsson, Genotypic Differences in Nitrate Uptake and Nitrogen Utilization for Spring Wheat Grown Hydroponically, Crop Science, vol.35, issue.4, pp.1056-1062, 1995.
DOI : 10.2135/cropsci1995.0011183X003500040023x

L. Peterson, G. Kleinkopf, and R. Huffaker, Evidence for Lack of Turnover of Ribulose 1,5-Diphosphate Carboxylase in Barley Leaves, PLANT PHYSIOLOGY, vol.51, issue.6, pp.1042-1045, 1973.
DOI : 10.1104/pp.51.6.1042

T. Pons and Y. De-jong-van-berkel, Species-specific Variation in the Importance of the Spectral Quality Gradient in Canopies as a Signal for Photosynthetic Resource Partitioning, Annals of Botany, vol.94, issue.5, pp.725-732, 2004.
DOI : 10.1093/aob/mch197

T. Pons, W. Jordi, and D. Kuiper, Acclimation of plants to light gradients in leaf canopies: evidence for a possible role for cytokinins transported in the transpiration stream, Journal of Experimental Botany, vol.52, issue.360, pp.1563-1574, 2001.
DOI : 10.1093/jexbot/52.360.1563

T. Pons, H. Van-rijnbeck, I. Scheurwater, . Van-der, and A. Verf, Importance of the gradient in photosynthetically active radiation in a vegetation stand for leaf nitrogen allocation in two monocotyledons, Oecologia, vol.30, issue.3, pp.416-424, 1993.
DOI : 10.1007/BF00320997

J. Reynolds and J. Chen, Modelling whole-plant allocation in relation to carbon and nitrogen supply: Coordination versus optimization: Opinion, Plant and Soil, vol.75, issue.76, pp.65-74, 1996.
DOI : 10.1007/BF02257565

P. Robson, I. Donnison, K. Wang, B. Frame, S. Pegg et al., Leaf senescence is delayed in maize expressing the Agrobacterium IPT gene under the control of a novel maize senescence-enhanced promoter, Plant Biotechnology Journal, vol.50, issue.2, pp.101-112, 2004.
DOI : 10.1046/j.1467-7652.2004.00054.x

M. Rousseaux, A. Hall, and R. Sanchez, Light Environment, Nitrogen Content, and Carbon Balance of Basal Leaves of Sunflower Canopies, Crop Science, vol.39, issue.4, pp.1093-1100, 1999.
DOI : 10.2135/cropsci1999.0011183X003900040023x

V. Sadras, L. Echarte, and F. Andrade, Profiles of Leaf Senescence During Reproductive Growth of Sunflower and Maize, Annals of Botany, vol.85, issue.2, pp.187-195, 2000.
DOI : 10.1006/anbo.1999.1013

F. Schieving, M. Werger, and T. Hirose, Canopy structure, nitrogen distribution and whole canopy photosynthetic carbon gain in growing and flowering stands of tall herbs, Vegetatio, vol.6, issue.2, pp.173-181, 1992.
DOI : 10.1007/BF00044732

V. Shearman, R. Sylvester-bradley, R. Scott, and M. Foulkes, Physiological processes associated with wheat yield progress in the UK, Crop Sci, vol.45, pp.175-185, 2005.

R. Simpson, H. Lambers, and M. Dalling, Nitrogen Redistribution during Grain Growth in Wheat (Triticum aestivum L.) : IV. Development of a Quantitative Model of the Translocation of Nitrogen to the Grain, PLANT PHYSIOLOGY, vol.71, issue.1, pp.7-14, 1983.
DOI : 10.1104/pp.71.1.7

R. Austin, Yield of Wheat in the United Kingdom, Crop Science, vol.39, issue.6, pp.1604-1610, 1999.
DOI : 10.2135/cropsci1999.3961604x

A. Barneix and M. Guitman, Leaf Regulation of the Nitrogen Concentration in the Grain of Wheat Plants, Journal of Experimental Botany, vol.44, issue.10, 1993.
DOI : 10.1093/jxb/44.10.1607

J. Bertheloot, P. Martre, and B. Andrieu, Dynamics of light and nitrogen distribution during grain filling within wheat canopy. Plant Physiology, in press. doi: 10, p.124156, 1104.
URL : https://hal.archives-ouvertes.fr/hal-00964201

A. Borrell and G. Hammer, Nitrogen Dynamics and the Physiological Basis of Stay-Green in Sorghum, Crop Science, vol.40, issue.5, pp.1295-1307, 2000.
DOI : 10.2135/cropsci2000.4051295x

M. Chelle, Phylloclimate or the climate perceived by individual plant organs: What is it? How to model it? What for? The New Phytologist 166, pp.781-790, 2005.

M. Chelle and B. Andrieu, The nested radiosity model for the distribution of light within plant canopies, Ecological Modelling, vol.111, issue.1, pp.75-91, 1998.
DOI : 10.1016/S0304-3800(98)00100-8

J. Chen, J. Reynolds, P. Harley, and J. Tenhunen, Coordination theory of leaf nitrogen distribution in a canopy, Oecologia, vol.39, issue.1, pp.63-69, 1993.
DOI : 10.1007/BF00321192

H. Cooper and D. Clarkson, Cycling of Amino-Nitrogen and other Nutrients between Shoots and Roots in Cereals???A Possible Mechanism Integrating Shoot and Root in the Regulation of Nutrient Uptake, Journal of Experimental Botany, vol.40, issue.7, pp.753-762, 1989.
DOI : 10.1093/jxb/40.7.753

M. Dreccer, G. Slafer, and R. Rabbinge, Optimization of Vertical Distribution of Canopy Nitrogen, Journal of Crop Production, vol.57, issue.1, pp.47-77, 1998.
DOI : 10.1016/0378-4290(93)90144-C

J. Evans, J. Evers, J. Vos, C. Fournier, B. Andrieu et al., Photosynthesis and nitrogen relationships in leaves of C3 plants, Oecologia, vol.282, issue.1, pp.9-19, 1989.
DOI : 10.1007/BF00377192

T. Evers and S. Millar, Cereal Grain Structure and Development: Some Implications for Quality, Journal of Cereal Science, vol.36, issue.3, p.435, 2002.
DOI : 10.1006/jcrs.2002.0435

C. Fournier, B. Andrieu, S. Ljutovac, S. Saint-jean, M. Hu et al., ADEL-wheat: a 3D architectural model of wheat development. In 'Plant Growth Modeling and Applications, Proceedings of the 2003 International Symposium, pp.54-63, 2003.
URL : https://hal.archives-ouvertes.fr/hal-00909184

C. Foyer, S. Ferrario-mery, and G. Noctor, Interactions Between Carbon and Nitrogen Metabolism, pp.237-244, 2001.
DOI : 10.1007/978-3-662-04064-5_9

O. Franklin and G. Agren, Leaf senescence and resorption as mechanisms of maximizing photosynthetic production during canopy development at N limitation, Functional Ecology, vol.102, issue.6, pp.727-733, 2002.
DOI : 10.1046/j.1469-8137.1999.00431.x

M. Girard, Modélisation de l'accumulation de biomasse et d'azote dans les grains de blé tendre d'hiver (Triticum aestivum L.); simulation de leur teneur en protéines à la récolte, 1997.

R. Gleadow, M. Dalling, and G. Halloran, Variation in Endosperm Characteristics and Nitrogen Content in Six Wheat Lines, Australian Journal of Plant Physiology, vol.9, issue.5, pp.539-551, 1982.
DOI : 10.1071/PP9820539

R. Graybosch, C. Peterson, D. Shelton, and P. Baenziger, Genotypic and Environmental Modification of Wheat Flour Protein Composition in Relation to End-Use Quality, Crop Science, vol.36, issue.2, pp.296-300, 1996.
DOI : 10.2135/cropsci1996.0011183X003600020014x

T. Hirose, Development of the Monsi-Saeki Theory on Canopy Structure and Function, Annals of Botany, vol.95, issue.3, pp.483-494, 2005.
DOI : 10.1093/aob/mci047

L. Irving and D. Robinson, A dynamic model of Rubisco turnover in cereal leaves, New Phytologist, vol.162, issue.3, 2006.
DOI : 10.1111/j.1469-8137.2005.01584.x

P. Jamieson and M. Semenov, Modelling nitrogen uptake and redistribution in wheat, Field Crops Research, vol.68, issue.1, pp.21-29, 2000.
DOI : 10.1016/S0378-4290(00)00103-9

R. Karwowski and P. Prusinkiewicz, Design and Implementation of the L+C Modeling Language, Electronic Notes in Theoretical Computer Science, vol.86, issue.2, pp.134-152, 2003.
DOI : 10.1016/S1571-0661(04)80680-7

O. Kull, Acclimation of photosynthesis in canopies: models and limitations, Oecologia, vol.133, issue.3, pp.267-279, 2002.
DOI : 10.1007/s00442-002-1042-1

O. Kull and B. Kruijt, Acclimation of photosynthesis to light: a mechanistic approach, Functional Ecology, vol.13, issue.1, 1999.
DOI : 10.1146/annurev.arplant.40.1.119

D. Lawlor, Carbon and nitrogen assimilation in relation to yield: mechanisms are the key to understanding production systems, Journal of Experimental Botany, vol.53370, pp.773-787, 2002.

M. Lötscher, K. Stroh, and H. Schnyder, Vertical Leaf Nitrogen Distribution in Relation to Nitrogen Status in Grassland Plants, Annals of Botany, vol.92, issue.5, pp.679-688, 2003.
DOI : 10.1093/aob/mcg188

C. Mackown, V. Sanford, D. Zhang, and N. , Wheat Vegetative Nitrogen Compositional Changes in Response to Reduced Reproductive Sink Strength, PLANT PHYSIOLOGY, vol.99, issue.4, pp.1469-1474, 1992.
DOI : 10.1104/pp.99.4.1469

D. Makowski, J. Hillier, D. Wallach, B. Andrieu, J. Mheds et al., Parameter estimation for crop modelsWorking with dynamic crop models. Evaluation, analysis, parameterisation and applications, pp.101-149, 2006.

P. Martre, J. Porter, P. Jamieson, and E. Triboï, Modeling Grain Nitrogen Accumulation and Protein Composition to Understand the Sink/Source Regulations of Nitrogen Remobilization for Wheat, PLANT PHYSIOLOGY, vol.133, issue.4, pp.1959-1967, 2003.
DOI : 10.1104/pp.103.030585

P. Martre, P. Jamieson, M. Semenov, R. Zyskowski, J. Porter et al., Modelling protein content and composition in relation to crop nitrogen dynamics for wheat, European Journal of Agronomy, vol.25, issue.2, 2006.
DOI : 10.1016/j.eja.2006.04.007

C. Masclaux, I. Quillere, A. Gallais, and B. Hirel, The challenge of remobilisation in plant nitrogen economy. A survey of physioagronomic and molecular approaches, The Annals of Applied Biology, vol.138, 2001.

M. Monsi and T. Saeki, On the Factor Light in Plant Communities and its Importance for Matter Production, Annals of Botany, vol.95, issue.3, pp.549-567, 2001.
DOI : 10.1093/aob/mci052

P. Prusinkiewicz, R. Karwowski, R. Mech, J. Hanan, and R. Mech, L-Studio/cpfg: A Software System for Modeling Plants, Lecture Notes in Computer Science, vol.1779, pp.457-464, 2000.
DOI : 10.1007/3-540-45104-8_38

J. Reynolds, J. Chen, V. Sadras, A. Hall, and D. Connor, Modelling whole-plant allocation in relation to carbon and nitrogen supply: Coordination versus optimization: Opinion, Plant and Soil, vol.75, issue.76, pp.65-74, 1993.
DOI : 10.1007/BF02257565

F. Schieving, M. Werger, T. Hirose, V. Shearman, R. Sylvester-bradley et al., Canopy structure, nitrogen distribution and whole canopy photosynthetic carbon gain in growing and flowering stands of tall herbs Physiological processes associated with wheat yield progress in the UK, Crop Science, vol.102, issue.45, pp.173-181, 1992.

P. Shewry, Improving the protein content and composition of cereal grain, Journal of Cereal Science, vol.46, issue.3, 2007.
DOI : 10.1016/j.jcs.2007.06.006

H. Shiratsuchi, T. Yamagishi, and R. Ishii, Leaf nitrogen distribution to maximize the canopy photosynthesis in rice, Field Crops Research, vol.95, issue.2-3, pp.291-304, 2006.
DOI : 10.1016/j.fcr.2005.04.005

R. Simpson, H. Lambers, and M. Dalling, Nitrogen redistribution during grain growth in wheat (Triticum aestivum L.), Planta, vol.11, issue.5, pp.7-14, 1983.
DOI : 10.1007/BF00386538

T. Sinclair, D. Wit, and C. , Photosynthate and Nitrogen Requirements for Seed Production by Various Crops, Science, vol.189, issue.4202, pp.565-567, 1975.
DOI : 10.1126/science.189.4202.565

B. Singh and C. Jenner, A modified method for the determination of cell number in wheat endosperm, Plant Science Letters, vol.26, issue.2-3, pp.273-278, 1982.
DOI : 10.1016/0304-4211(82)90101-8

J. Soussana, F. Teyssonneyre, and J. Thierry, Un modèle dynamique d'allocation basé sur l'hypothèse d'une colimitation de la croissance végétale par les absorptions de lumière et d'azote. In 'Fonctionnement des peuplements végétaux sous contraintes environnementales, pp.87-116, 2000.

F. Tabourel-tayot and F. Gastal, MecaNiCAL, a supply???demand model of carbon and nitrogen partitioning applied to defoliated grass, European Journal of Agronomy, vol.9, issue.4, pp.223-241, 1998.
DOI : 10.1016/S1161-0301(98)00039-2

E. Tambussi, J. Bort, J. Guiamet, S. Nogués, and J. Araus, Cereals: Metabolism, Water Use Efficiency and Contribution to Grain Yield, Critical Reviews in Plant Sciences, vol.59, issue.1, pp.1-16, 2007.
DOI : 10.1007/BF00393191

H. Thomas, H. Ougham, C. Wagstaff, and A. Stead, Defining senescence and death, Journal of Experimental Botany, vol.54, issue.385, 2003.
DOI : 10.1093/jxb/erg133
URL : http://citeseerx.ist.psu.edu/viewdoc/summary?doi=10.1.1.320.1894

J. Thornley, Dynamic Model of Leaf Photosynthesis with Acclimation to Light and Nitrogen, Annals of Botany, vol.81, issue.3, pp.421-430, 1997.
DOI : 10.1006/anbo.1997.0575

J. Thornley, Acclimation of Photosynthesis to Light and Canopy Nitrogen Distribution: an Interpretation, Annals of Botany, vol.93, issue.4, pp.473-475, 2004.
DOI : 10.1093/aob/mch051

E. Triboï and A. Triboï-blondel, Productivity and grain or seed composition: a new approach to an old problem???invited paper, European Journal of Agronomy, vol.16, issue.3, pp.163-186, 2002.
DOI : 10.1016/S1161-0301(01)00146-0

E. Triboï, P. Martre, and A. Triboi-blondel, Environmentally-induced changes in protein composition in developing grains of wheat are related to changes in total protein content, Journal of Experimental Botany, vol.54, issue.388, pp.1731-1742, 2003.
DOI : 10.1093/jxb/erg183

X. Yin, H. Schapendonk, M. Kropff, M. Van-oijen, and P. Bindraban, A Generic Equation for Nitrogen-limited Leaf Area Index and its Application in Crop Growth Models for Predicting Leaf Senescence, Annals of Botany, vol.85, issue.5, pp.579-585, 2000.
DOI : 10.1006/anbo.1999.1104

D. Ackerly, Self-shading, carbon gain and leaf dynamics: a test of alternative optimality models, Oecologia, vol.119, issue.3, pp.300-310, 1999.
DOI : 10.1007/s004420050790

D. Ackerly and F. Bazzaz, Leaf dynamics, self-shading and carbon gain in seedlings of a tropical pioneer tree, Oecologia, vol.36, issue.3, pp.289-298, 1995.
DOI : 10.1007/BF00328814

B. Andrieu, J. Lecoeur, G. Lemaire, B. Ney, . Doré et al., Le peuplement végétal cultivé, J Roger-Estrade, pp.103-136, 2006.

M. Bancal, C. Robert, and B. Ney, Modelling Wheat Growth and Yield Losses from Late Epidemics of Foliar Diseases using Loss of Green Leaf Area per Layer and Pre-anthesis Reserves, Annals of Botany, vol.100, issue.4, pp.777-789, 2007.
DOI : 10.1093/aob/mcm163
URL : https://hal.archives-ouvertes.fr/hal-01191942

J. Bertheloot, P. Martre, and B. Andrieu, Dynamics of Light and Nitrogen Distribution during Grain Filling within Wheat Canopy, PLANT PHYSIOLOGY, vol.148, issue.3, pp.1707-1720, 2008.
DOI : 10.1104/pp.108.124156
URL : https://hal.archives-ouvertes.fr/hal-00964201

A. Borrell and G. Hammer, Nitrogen Dynamics and the Physiological Basis of Stay-Green in Sorghum, Crop Science, vol.40, issue.5, pp.1295-1307, 2000.
DOI : 10.2135/cropsci2000.4051295x

M. Chelle and B. Andrieu, The nested radiosity model for the distribution of light within plant canopies, Ecological Modelling, vol.111, issue.1, pp.75-91, 1998.
DOI : 10.1016/S0304-3800(98)00100-8

H. Cooper, D. Clarkson, M. Johnston, J. Whiteway, and B. Loughman, Cycling of amino-nitrogen between shoots and roots in wheat seedlings, Plant and Soil, p.91, 1986.

J. Evans, Photosynthesis and nitrogen relationships in leaves of C3 plants, Oecologia, vol.282, issue.1, pp.9-19, 1989.
DOI : 10.1007/BF00377192

J. Evans and J. Seemann, The allocation of protein nitrogen in the photosynthetic apparatus: costs, consequences and control, pp.183-205, 1989.

J. Evers, J. Vos, C. Fournier, B. Andrieu, M. Chelle et al., Towards a generic architectural model of tillering in Gramineae, as exemplified by spring wheat (Triticum aestivum), New Phytologist, vol.31, issue.3, pp.801-812, 2005.
DOI : 10.1111/j.1469-8137.2005.01337.x

C. Field, Allocating leaf nitrogen for the maximization of carbon gain: Leaf age as a control on the allocation program, Oecologia, vol.38, issue.2-3, pp.341-347, 1983.
DOI : 10.1007/BF00379710

C. Fournier and B. Andrieu, A 3D Architectural and Process-based Model of Maize Development, Annals of Botany, vol.81, issue.2, pp.233-250, 1998.
DOI : 10.1006/anbo.1997.0549

M. Guitman, P. Arnozis, and A. Barneix, Effect of source-sink relations and nitrogen nutrition on senescence and N remobilization in the flag leaf of wheat, Physiologia Plantarum, vol.82, 1991.

L. Irving and D. Robinson, A dynamic model of Rubisco turnover in cereal leaves, New Phytologist, vol.162, issue.3, pp.493-504, 2006.
DOI : 10.1111/j.1469-8137.2005.01584.x

P. Jamieson, M. Semenov, I. Brooking, and G. Francis, Sirius: a mechanistic model of wheat response to environmental variation, European Journal of Agronomy, vol.8, issue.3-4, pp.161-179, 1998.
DOI : 10.1016/S1161-0301(98)00020-3

M. Jeuffroy, B. Ney, and A. Ourry, Integrated physiological and agronomic modelling of N capture and use within the plant, Journal of Experimental Botany, vol.53, issue.370, pp.809-823, 2002.
DOI : 10.1093/jexbot/53.370.809

P. Martre, J. Porter, P. Jamieson, and E. , Modeling Grain Nitrogen Accumulation and Protein Composition to Understand the Sink/Source Regulations of Nitrogen Remobilization for Wheat, PLANT PHYSIOLOGY, vol.133, issue.4, pp.1959-1967, 2003.
DOI : 10.1104/pp.103.030585

H. Mooney, C. Field, S. Gulmon, and F. Bazzaz, Photosynthetic capacity in relation to leaf position in desert versus old-field annuals, Oecologia, vol.64, issue.1, pp.109-112, 1981.
DOI : 10.1007/BF00378802

J. Müller, P. Wernecke, and W. Diepenbrock, LEAFC3-N: a nitrogen-sensitive extension of the CO2 and H2O gas exchange model LEAFC3 parameterised and tested for winter wheat (Triticum aestivum L.), Ecological Modelling, vol.183, issue.2-3, pp.183-210, 2005.
DOI : 10.1016/j.ecolmodel.2004.07.025

T. Pons, J. De, and Y. Berkel, Species-specific Variation in the Importance of the Spectral Quality Gradient in Canopies as a Signal for Photosynthetic Resource Partitioning, Annals of Botany, vol.94, issue.5, pp.725-732, 2004.
DOI : 10.1093/aob/mch197

C. Robert, M. Bancal, and C. Lannou, Wheat Leaf Rust Uredospore Production and Carbon and Nitrogen Export in Relation to Lesion Size and Density, Phytopathology, vol.92, issue.7, pp.762-768, 2002.
DOI : 10.1094/PHYTO.2002.92.7.762

C. Robert, C. Fournier, B. Andrieu, and B. Ney, epidemic model (Septo3D): a new approach to investigate plant???pathogen interactions linked to canopy architecture, Functional Plant Biology, vol.35, issue.10, pp.997-1013, 2007.
DOI : 10.1071/FP08066
URL : https://hal.archives-ouvertes.fr/hal-01192058

C. Ta and R. Weiland, Nitrogen Partitioning in Maize during Ear Development, Crop Science, vol.32, issue.2, pp.443-451, 1992.
DOI : 10.2135/cropsci1992.0011183X003200020032x

E. Tambussi, J. Bort, J. Guiamet, S. Nogues, and J. Araus, Cereals: Metabolism, Water Use Efficiency and Contribution to Grain Yield, Critical Reviews in Plant Sciences, vol.59, issue.1, pp.1-16, 2007.
DOI : 10.1007/BF00393191

J. Thornley, Dynamic Model of Leaf Photosynthesis with Acclimation to Light and Nitrogen, Annals of Botany, vol.81, issue.3, pp.421-430, 1998.
DOI : 10.1006/anbo.1997.0575

J. Thornley, Acclimation of Photosynthesis to Light and Canopy Nitrogen Distribution: an Interpretation, Annals of Botany, vol.93, issue.4, pp.473-475, 2004.
DOI : 10.1093/aob/mch051

E. Triboi, P. Martre, and A. Triboi-blondel, Environmentally-induced changes in protein composition in developing grains of wheat are related to changes in total protein content, Journal of Experimental Botany, vol.54, issue.388, pp.1731-1742, 2003.
DOI : 10.1093/jxb/erg183