M. Ames, D. And, and . Spooner, DNA from herbarium specimens settles a controversy about origins of the European potato, American Journal of Botany, vol.95, issue.2, pp.252-257, 2008.
DOI : 10.3732/ajb.95.2.252

J. E. Bradshaw, G. J. Bryan, and G. Ramsay, genetic Resources (Including wild and cultivated Solanum Species) and progress in their utilization in potato breeding. Potato Research, pp.49-65, 2006.

A. Castronovo, Papas chilotas, 1949.

M. Ghislain, J. Núñez, M. Del, R. Herrera, and D. M. Spooner, The single Andigenum origin of Neo-Tuberosum potato materials is not supported by microsatellite and plastid marker analyses, Theoretical and Applied Genetics, vol.104, issue.5, pp.963-969, 2009.
DOI : 10.1007/s00122-008-0953-6

D. Glendinning, Chilean potatoes: an appraisal. Potato Research, pp.306-307, 1975.

. Greenpeace, Centros de diversidad. La riqueza biológica de los cultivos tradicionales, herencia mundial amenazada por la contaminación genética, 2000.

P. Grun, Evoluti??n de Papas Cultivadas, Economic Botany, vol.6, issue.S3, pp.39-55, 1990.
DOI : 10.1007/BF02860474

J. G. Hawkes, The potato. Evolution, biodiversity and genetic resources, p.178, 1990.

J. G. Hawkes and J. F. Ortega, La patata en Espa??a a finales del siglo XVI, Economic Botany, vol.161, issue.1, pp.86-97, 1992.
DOI : 10.1007/BF02985257

J. G. Hawkes and J. F. Ortega, The early history of the potato in Europe, Euphytica, vol.46, issue.1, pp.1-7, 1993.
DOI : 10.1007/BF00029633

J. Hawkes, Origins of cultivated potatoes and species relatioships Potato Genetics, pp.3-39, 1994.

K. Hosaka, G. A. De-zoeten, R. E. Hanneman, and J. , Cultivated potato chloroplast DNA differs from the wild type by one deletion ??? evidence and implications, Theoretical and Applied Genetics, vol.75, issue.5, 1988.
DOI : 10.1007/BF00265598

K. Hosaka, Succesive domestication and evolution of the Andean potatoes as revealed by chloroplast DNA restriction endonuclease analysis, Theor. Appl. Genet, vol.90, pp.356-363, 1995.

Z. Huamán and D. M. Spooner, Reclassification of landrace populations of cultivated potatoes (Solanum sect. Petota), American Journal of Botany, vol.89, issue.6, pp.947-965, 2002.
DOI : 10.3732/ajb.89.6.947

S. H. Jansky and S. J. Peloquin, Advantages of Wild Diploid Solanum Species Over Cultivated Diploid Relatives in Potato Breeding Programs, Genetic Resources and Crop Evolution, vol.66, issue.3 supplement, pp.669-674, 2006.
DOI : 10.1007/s10722-004-2949-7

Y. Kawagoe and K. , Chloroplast DNA evolution in potato (Solanum tuberosum L.), Theoretical and Applied Genetics, vol.81, issue.1, pp.13-20, 1991.
DOI : 10.1007/BF00226106

A. Lössl, N. Adler, R. Horn, U. Frei, and G. Wenzel, Chondriome-type characterization of potato: mt ? ,? , ?, ?, ? and novel plastid-mitochondrial configurations in somatic hybrids, Theor. Appl. Genet, pp.981-991, 1999.

A. Montaldo, Cultivo y mejoramiento de la papa. Instituto Interamericano de Cooperación para la Agricultura, 1984.

C. Raker and D. Spooner, Chilean Tetraploid Cultivated Potato, , is Distinct from the Andean Populations, Crop Science, vol.42, issue.5, pp.1451-1458, 2002.
DOI : 10.2135/cropsci2002.1451

D. Rios, M. Ghislain, F. Rodriguez, D. And, and . Spooner, What Is the Origin of the European Potato? Evidence from Canary Island Landraces, Crop Science, vol.47, issue.3, pp.1271-1280, 2007.
DOI : 10.2135/cropsci2006.05.0336

J. L. Ruiz-de-galarreta, J. Pascualena, F. J. Legorburu, L. Barandalla, and E. Ritter, The History of Potato (Solanum tuberosum L.) Research in Spain, Potato Research, vol.14, issue.1, pp.19-25, 2006.
DOI : 10.1007/s11540-006-9001-6

N. W. Simmonds, Potatoes ? Solanum tuberosum (Solanaceae), In: J. Smartt and N, 1995.

D. M. Spooner and W. L. Hetterscheid, Origins, evolution, and group classification of cultivated potatoes. P.285-307 Darwin´s harvest: New approaches to the origins, evolution and conservation of crops, 2005.

. D. Spooner, . Mc, K. Lean, G. Ramsay, R. Waugh et al., A single domestication for potato based on multilocus amplified fragment length polymorphism genotyping, Proceedings of the National Academy Sciences of the United States of America, pp.14694-14699, 2005.
DOI : 10.1073/pnas.0507400102

J. D. Spooner, F. Nuñez, P. Rodriguez, M. Nair, and . Ghislain, Nuclear and chloroplast DNA reassessment of the origin of Indian potato varieties and its implications for the origin of the early European potato, Theoretical and Applied Genetics, vol.66, issue.Suppl 3, pp.1020-1026, 2004.
DOI : 10.1007/s00122-004-1917-0

D. M. Spooner, J. Nuñez, G. Trujillo, M. Herrera, F. Guzman et al., Extensive simple sequence repeat genotyping of potato landraces supports a major reevaluation of their gene pool structure and classification, Proc. Natl. Acad. Sci, pp.19398-10403, 2007.
DOI : 10.1073/pnas.0709796104

R. G. Van-den-berg, J. T. Miller, M. L. Ugarte, J. P. Kardolus, J. Nienhuis et al., Collapse of Morphological Species in the Wild Potato Solanum brevicaule Complex (Solanaceae: Sect. Petota), American Journal of Botany, vol.85, issue.1, pp.92-109, 1998.
DOI : 10.2307/2446559

E. M. Boeninger, H. Franco, G. Ortega, C. Irarrazabal, O. Valdes et al., La economía campesina, 1981.

S. B. Brush, Ethnoecology, biodiversity and modernization in Andean potato agriculture, Journal of Ethnobiology, vol.12, pp.161-185, 1992.

R. Cárdenas, La papa, patrimonio de la humanidad. Revista de la papa, N°, vol.5, issue.15, pp.14-15, 2003.

M. Cox, Agricultura chilena 1974-1982, 1983.

A. Cubillos, Chile: Informe nacional para la conferencia técnica internacional de la FAO sobre los recursos filogenéticos, 1995.

;. M. Tanto, . Sawadogo-;-i, ;. Mar, . Sadiki-;-n, ;. L. Thi-ngoc-hue et al., Bajracharya; F. Castillo, Jwala Deepak Rijal; L. Belqadik; R. Rana; S

J. Ouedraogo-;-r, ;. L. Zangre, ;. Chavez, . Schoen-;-b, ;. Sthapit et al., A global perspective of the richness and evenness of traditional crop-variety diversity maintained by farming communities, 2008.

C. Gay, Historia física y política de Chile, Paris. MD CCCLXV. Francia, p.443, 1974.

S. Gómez, Instituciones y procesos agrarios en Chile. FLACSO, 1982.

J. Hair, R. Anderson, R. Tatham, and W. Black, Análisis Multivariante, 1999.

R. J. Hijmans and D. M. Spooner, Geographic Distribution of Wild Potato Species, American Journal of Botany, vol.88, issue.11, 2001.
DOI : 10.2307/3558435

J. Kalazich, Variedades aptas para el mercado. Tierra Adentro, 2008.

M. De-agricultura, Ley-18919, 1990.

M. De and A. , Políticas y bases programáticas del desarrollo de la pequeña agricultura campesina, la reducción de la pobreza rural y el fortalecimiento de las organizaciones de productores, Proyecto TCP, vol.23, p.51, 1992.

M. De and A. , Una política de estado para la agricultura chilena, 2000.

A. Montaldo, Cultivo y mejoramiento de la papa. Instituto Interamericano de Cooperación para la Agricultura, pp.359-414, 1984.

V. Brush and S. B. , Ethnoecology, biodiversity and modernization in Andean potato agriculture, Journal of Ethnobiology, vol.12, pp.161-185, 1992.

P. Dacom-plant, Plant plus manual Using plant-plus on the internet, 2003.

C. Dervin, Comment interpréter les résultats d´une analyse factorielle des correspondances, Publication ITCF. Institut Technique des Céréales et des Fourrages, p.75, 1990.

C. Dervin, Comment interpréter les résultats d´une classification automatique. Publication ITCF. Coleccection STAT-ITCF. ITCF Céréaliers De France, p.70, 1996.

. J. Hawkes, The potato. Evolution, biodiversity and genetic resources, p.178, 1990.

J. Z. Huaman, W. Williams, N. Salhuana, and . Vincent, Descriptors for the cultivated potato and for the maintenance and distribution of germoplasm collections, 1977.

C. B. Quiros, D. Brush, K. Douches, G. Zimmerer, and . Huestis, Evaluati??n bioqu??mica y popular de la variabilidad interespec??fica de las papas cultivadas en los Andes, Economic Botany, vol.20, issue.2, pp.254-266, 1990.
DOI : 10.1007/BF02860490

K. D. Spooner, G. Mclean, R. Ramsay, G. Waug, and . Bryan, A single domestication for potato based on multilocus amplified fragment length polymorphism genotyping, Proceeding of the National Academy of Sciences, pp.14694-14699, 2005.
DOI : 10.1073/pnas.0507400102

D. M. Ghislain, F. Andrade, R. Rodríguez, D. Hijmans, and . Spooner, Genetic analysis of the cultivated potato Solanum tuberosum L. Phureja Group using RAPDs and nuclear SSRs, Theoretical and Applied Genetics, vol.18, issue.8, pp.1515-1527, 2006.
DOI : 10.1007/s00122-006-0399-7

J. M. Ghislain, . M. Nuñez, R. Del, D. Herrera, and . Spooner, The single Andigenum origin of Neo-Tuberosum potato materials is not supported by microsatellite and plastid marker analyses, Theoretical and Applied Genetics, vol.104, issue.5, pp.693-969, 2009.
DOI : 10.1007/s00122-008-0953-6

. Spooner, Robust and highly informative microsatellite-based genetic identity kit for potato, Mol. Breeding, vol.23, pp.377-388, 2009.

Y. Gong, S. Xu, W. Mao, Q. Hu, G. Zhang et al., Developing new SSR markers from ESTs of pea (Pisum sativum L.), Journal of Zhejiang University SCIENCE B, vol.11, issue.9, pp.702-707, 2010.
DOI : 10.1631/jzus.B1000004

R. J. Hijmans and D. M. Spooner, Geographic Distribution of Wild Potato Species, American Journal of Botany, vol.88, issue.11, pp.2101-2112, 2001.
DOI : 10.2307/3558435

S. Hurlbert, The Nonconcept of Species Diversity: A Critique and Alternative Parameters, Ecology, vol.52, issue.4, pp.577-586, 1971.
DOI : 10.2307/1934145

V. Ispizúa, I. Guma, S. Feingold, and A. Clausen, Genetic diversity of potato landraces from northwestern Argentina assessed with simple sequence repeats (SSRs) Genetic Resources Crop Evolution, pp.1833-1848, 2007.

G. Levinson and G. A. Gutman, Slipped-strand mispairing: A major mechanism for DNA sequence evolution, Mol. Biol. Evol, pp.203-221, 1987.

P. Marchezi, T. De-campos, A. Barbosa-de-sousa, D. Sforca, G. Montan et al., Potato cultivar identification using molecular markers, 2010.

M. Mathias, B. Sagredo, and K. Kalazich, SSR molecular markers for identification of potato germplasm in the breeding program at INIA in Chile, Agricultura Técnica (Chile), vol.67, issue.1, pp.3-15, 2007.

L. P. Martinez, . Cavagnaro-;-r, J. Masuelli, and . Rodriguez, Evaluation of diversity among Argentine grapevine (Vitis vinifera L.) varietes using morphological data and AFLP markers, Electronical Journal Biotechnology, vol.6, issue.3, pp.241-250, 2003.

C. Mcgregor, M. Greyling, C. Lambert, J. Louw, and L. Warnich, A comparative assessment of DNA fingerprinting techniques (RAPS, ISSR, AFLP and SSR) in tetraploid potato (Solanum tuberosum L.) germplasm, Euphytica, vol.113, issue.2, pp.135-144, 2000.
DOI : 10.1023/A:1003925620546

D. Milbourne, R. Meyer, A. Collins, L. Ramsay, C. Gebhardt et al., Isolation, characterisation and mapping of simple sequence repeat loci in potato, Molecular and General Genetics MGG, vol.259, issue.3, pp.233-245, 1988.
DOI : 10.1007/s004380050809

J. T. Miller and D. Spooner, Collapse of species boundaries in the wild potatoSolanum brevicaule complex (Solanaceae, S. sect.Petota): Molecular data, Plant Systematics and Evolution, vol.36, issue.1-4, pp.103-130, 1999.
DOI : 10.1007/BF00985734

S. A. Mohammadi and B. M. Prasanna, Analysis of Genetic Diversity in Crop Plants???Salient Statistical Tools and Considerations, Crop Science, vol.43, issue.4, pp.1235-1248, 2003.
DOI : 10.2135/cropsci2003.1235

M. Moisan-thiersy, S. Marhadour, M. Kerlan, N. Dessenne, M. Perramant et al., Potato cultivar identification using simple sequence repeats markers (SSR), Potato Research, vol.31, issue.1, pp.191-200, 2005.
DOI : 10.1007/BF02742376

K. B. Mullis and F. Faloona, Specific synthesis of DNA in vitro via polymerase chain reaction, Methods Enzymol, vol.155, pp.350-355, 1987.

. M. Nei, Analysis of Gene Diversity in Subdivided Populations, Proc. Natl. Acad. Sci. USA. Vo, pp.3321-3323, 1973.
DOI : 10.1073/pnas.70.12.3321

R. Ortiz and Z. Huaman, Inheritance of morphological and tuber characteristics, In: CAB International. Potato Genetics, pp.263-279, 1994.

J. Ortiz, I. Aguimagalde, and J. Martín, Varietal identification. P. 515-560 Los marcadores genéticos en la mejora vegetal, 2000.

X. Pierrer and J. P. Jacquemoud-collet, DARwin software, 2006.

C. M. Raker and D. M. Spooner, Chilean Tetraploid Cultivated Potato, , is Distinct from the Andean Populations, Crop Science, vol.42, issue.5, pp.1451-1458, 2002.
DOI : 10.2135/cropsci2002.1451

A. Reid and E. M. Kerr, A rapid simple sequence repeat (SSR)-based identification method for potato cultivars, Plant Genetic Resources: Characterization and Utilization, vol.70, issue.01, pp.7-13, 2007.
DOI : 10.1007/BF02883885

A. Ritter, Aplicación de la biotecnología a la mejora genética de la patata, PATATA 2000, p.8, 2000.

I. F. Roldán-ruiz, ;. T. Van-eeuwijk, P. Gilliland, C. Dubreil, ;. J. Dillman et al., A comparative study of molecular and morphological methods of describing relationships between perennial ryegrass ( Lolium perenne L.) varieties, TAG Theoretical and Applied Genetics, vol.103, issue.8, pp.1138-1150
DOI : 10.1007/s001220100571

S. Maroof, M. A. , R. M. Biyashev, G. P. Yang, Q. Zhang et al., Extraordinarily polymorphic microsatellite DNA in barley: species diversity, chromosomal locations, and population dynamics, Proc. Natl. Acad. Sci. USA, pp.5466-5470, 1994.

N. Saitou and M. Nei, The Neighbor-Joining method: a new method for reconstructing phylogenetic trees, Molecular Biology and Evolution, vol.4, issue.4, pp.406-425, 1987.

C. Schlotterer and D. Tautz, Slippage synthesis of simple sequence DNA, Nucleic Acids Research, vol.20, issue.2, pp.2211-2215, 1992.
DOI : 10.1093/nar/20.2.211

D. M. Spooner, I. Peralta, and S. Knapp, Comparison of AFLPs with Other Markers for Phylogenetic Inference in Wild Tomatoes [Solanum L. Section Lycopersicon (Mill.) Wettst.], Taxon, vol.54, issue.1, 2005.
DOI : 10.2307/25065301

. Wettst, Taxon, pp.43-61

D. Spooner, K. Mclean, G. Ramsay, R. Waugh, and G. Bryan, A single domestication for potato based on multilocus amplified fragment length polymorphism genotyping, Proceeding of the National Academy of Sciences, pp.14694-14699, 2005.
DOI : 10.1073/pnas.0507400102

D. Spooner, J. Nuñez, G. Trujillo, R. Herrera, F. Guzman et al., Extensive simple sequence repeat genotyping of potato landraces supports a major reevaluation of their gene pool structure and classification, Proceedings of the National Academy of Sciences, vol.104, issue.49, pp.19398-19403, 2007.
DOI : 10.1073/pnas.0709796104

H. Stegemann and V. Loeschecke, Index. Europäischer Kartoffelsorten; Bestimmung durch elecktrophoretische Spektren, p.214, 1976.

G. Sun, G. Wang-pruski, M. Mayich, and H. Jong, RAPD and pedigree-based genetic diversity estimates in cultivated diploid potato hybrids, Theoretical and Applied Genetics, vol.107, issue.1, pp.110-115, 2003.
DOI : 10.1007/s00122-003-1215-2

D. Tautz and M. Renz, Simple sequences are ubiquitous repetitive components of eukaryotic genomes, Nucleic Acids Research, vol.12, issue.10, pp.4127-4138, 1984.
DOI : 10.1093/nar/12.10.4127

C. Van-de-wiel, P. Arens, and B. Vosman, Microsatellite retrieval in lettuce (<i>Lactuca sativa</i> L.), Genome, vol.42, issue.1, pp.139-149, 1999.
DOI : 10.1139/gen-42-1-139

S. D. Wu and K. S. Tanksley, Abundance, polymorphism and genetic mapping of microsatellites in rice, MGG Molecular & General Genetics, vol.18, issue.1-2, pp.225-235, 1993.
DOI : 10.1007/BF00280220

I. , D. A. Taylor, P. W. Ford, R. Franz, P. Mc et al., DNA fingerprinting and genetic relationships of potato cultivars (Solanum tuberosum L.) commercially grown in Australia, Australian Journal of Agricultural Research, vol.52, pp.911-918, 2001.

K. J. Eck, B. R. Van, . Van, and . Den, The potencial of AFLPs in biosystematics: a first application in Solanum taxonomy (Solanaceae). Plant Systematicd and Evolution, pp.87-103, 1998.

K. , J. H. Joung, H. Kim, H. Y. , L. et al., Estimation of genetic variation and relationship in potato

M. , P. J. Saghai-maroof, M. A. Buss, G. R. , H. et al., Amplified fragment length polymorphism (AFLP) in soybean: species diversity, inheritance, and near-isogenic line analysis, Theoretical and Applied Genetics, vol.93, issue.3, pp.392-401, 1996.

M. , L. Cavagnaro, P. Masuelli, R. , R. et al., Evaluation of diversity among Argentine grapevine (Vitis vinifera L.) varietes using morphological data and AFLP markers, Electronical Journal Biotechnology, vol.6, issue.3, pp.241-250, 2003.

C. E. Mcgregor, M. M. Greyling, C. A. Lambert, J. H. Louw, W. et al., A comparative assessment of DNA fingerprinting techniques (RAPS, ISSR, AFLP and SSR) in tetraploid potato (Solanum tuberosum L.) germplasm, Euphytica, vol.113, issue.2, pp.135-144, 2000.
DOI : 10.1023/A:1003925620546

C. E. Mcgregor, R. Van-treuren, R. Hockstra, V. Hintum, and T. J. , Analysis of the wild potato germplasm of the series Acaulia with AFLPs: implications for ex situ conservation, TAG Theoretical and Applied Genetics, vol.104, issue.1, pp.146-156, 2002.
DOI : 10.1007/s001220200018

M. , D. Meyer, R. Collins, A. Ramsay, L. Gebhardt et al., Isolation, characterisation and mapping of simple sequence repeat loci in potato, Molecular and General Genetics, vol.259, issue.3, pp.233-245, 1998.

P. , S. Wachira, F. N. Powell, W. , W. et al., Diversity and genetic differentiation among populations of Indian and Kenyan tea (Camellia sinensis

P. , W. Morgante, M. Andre, C. Hanafey, M. Vogel et al., The comparison of RFLP, RAPD, AFLP and SSR (microsatellite) markers for germplasm analysis, Molecular Breeding, vol.2, pp.225-238, 1996.

R. , I. Van-eeuwijk, F. A. Gilliland, T. J. Dubreil, P. Dillman et al., A comparative study of molecular and morphological methods of describing relationships between perennial ryegrass (Lolium Perenne L.) varietes, Theoretical and Applied Genetics, vol.103, issue.8, pp.1138-1150, 2001.

S. , D. M. Peralta, I. , K. , and S. , Comparison of AFLPs with Other Markers for Phylogenetic Inference in Wild Tomatoes (Solanum L. Section Lycopersicon, Taxon, vol.54, pp.43-61, 2005.

S. , D. Mclean, K. Ramsay, G. Waugh, R. et al., A single domestication for potato based on multilocus amplified fragment length polymorphism genotyping, Proceeding of the National Academy of Sciences, pp.14694-14699, 2005.

S. , G. Wang-pruski, G. Mayich, M. , J. et al., RAPD and pedigree-based genetic diversity estimates in cultivated diploid potato hybrids, Theoretical and Applied Genetics, vol.107, issue.1, pp.110-115, 2003.

V. , P. Hogers, R. Bleeker, M. Reijans, M. Van-de-lee et al., AFLP: A new technique for DNA fingerprinting, Nucleic Acids Research, vol.23, issue.21, pp.4407-4414, 1995.

X. , R. Q. Tomooka, N. , V. , and D. A. , AFLP markers for characterizing the azuki bean complex, Crop Science, vol.40, pp.808-815, 2000.

Y. , E. Kidwell, K. K. Sills, G. R. , L. et al., Diversity among selected Vigna angularis (Azuki) accessions on the basis of RAPD and AFLP markers, Crop Science, vol.39, pp.268-275, 1999.

. Lucca, show that the variability existing in S. tuberosum ssp andigena (adg) and S. tarinjense have not yet been fully explored in the south east of Buenos Aires province in, 2008.

. Chiloé, The degrees of incidence and severity in some years reach 100% of the crop (Table 2). 100 Stems and leaves dead

I. Acuña, B. Sagredo, R. Bravo, M. Gutiérrez, I. Maldonado et al., Using a forecasting system to develop integrated pest management strategies for control of late blight in southern Chile. Proceeding of the tenth workshop of an European network for development of an integrated control strategy of potato late blight, pp.237-249, 2007.

I. Acuña, R. Bravo, J. Inostroza, N. Gaete, I. Maldonado et al., El tizón tardío en Chile. En : XXIII Congreso de la Asociación Latinoamericana de la papa y VI Seminario latinoamericano de Uso y Comercialización de la Papa: Memorias, 2008.

A. Andreu, D. Caldiz, and G. Forbes, Phenotypic Expression of Resistance to Phytophthora infestans in Processing Potatoes in Argentina, American Journal of Potato Research, vol.99, issue.2, pp.177-187, 2009.
DOI : 10.1007/s12230-009-9121-z

D. Andrivon, F. Pilet, J. Montarry, M. Hafidi, R. Corbiere et al., to Partial Resistance in Potato: Evidence from French and Moroccan Populations, Phytopathology, vol.97, issue.3, pp.338-343, 2007.
DOI : 10.1094/PHYTO-97-3-0338

D. Andrivon, J. Lucas, and . Ellisseche, Development of natural late blight epidemics in pure and mixed plots of potato cultivars with different levels of partial resistance, Plant Pathology, vol.64, issue.5, pp.586-594, 2003.
DOI : 10.1038/35021046

S. M. Al-kherb, C. Fininsa, R. C. Shattock, and D. S. Shaw, The inheritance of virulence of Phytophthora infestans to potato, Plant Pathology, vol.95, issue.3, pp.552-562, 1995.
DOI : 10.1016/0147-5975(87)90032-6

S. Arentsen, Fitopatología Chilena, Historia y Desarrollo Conference at VII Meeting of the Latinoamerican Phytopathological Society, 1994.

L. Barandalla, R. López, J. Ruiz-de-galarreta, and Y. E. Ritter, Evaluación frente a diferentes patógenos de una colección de cultivares antiguos y germoplasma nativo de papa, En: III Congreso Iberoamericano de Investigación y Desarrollo en Patata, 2008.

. Edicion, Administración de la Comunidad Autónoma del País Vasco Departamento de Agricultura, Pesca y Alimentación. 5-10 de Octubre, pp.59-62, 2008.

M. Barquero, L. Gómez, and A. Brenes, Resistencia al tizón tardío (Phytophthora infestans) en clones promisorios de papa en Costa Rica, Agronomía Costarricense, vol.29, issue.3, pp.31-45, 2005.

M. Barquero, L. Gómez, and A. Brenes, Resistencia al tizón tardío (Phytophthora infestans) en clones promisorios de papa en Costa Rica, Agronomía Costarricense, vol.29, issue.3, pp.31-45, 2005.

L. Ciampi, La patología vegetal en Chile, Revista Asociación Chilena de la Papa, vol.3, issue.9, pp.1-2, 2001.

L. Colon, D. Budding, L. Keizer, and M. Pieters, Components of resistance to late blight (Phytophthora infestans) in eight South AmericanSolanum species, European Journal of Plant Pathology, vol.77, issue.4, pp.441-456, 1995.
DOI : 10.1007/BF01874858

L. Colon, U. B. Nielsen, and . Darsow, EUCABLIGHT-Detached leaf test for foliage blight resistance, 2004.

M. Díaz, D. Fajardo, J. Dilmer, C. García, and V. Núñez, Identificación de genes R1 y R2 que confieren resistencia a Phytophthora infestans en genotipos colombianos de papa, Revista Colombiana de Biotecnología, vol.5, issue.2, pp.40-50, 2003.

M. Escalante and C. Armas, Evaluación epidemiológica y económica en el manejo del tizón tardío de la papa en las Playitas, Bioagro, vol.16, issue.3, pp.189-196, 2004.

R. N. Estrada, E. Fernández-northcote, E. , C. , and O. Navia, Mejoramiento genético para resistencia a enfermedades y plagas de la papa en Bolivia, Memorias del 1 er Taller Sobre Resistencia Duradera en Cultivos Alto Andinos de Bolivia, 1994.

W. Flier, L. Turkensteen, and A. Mulder, Variation in tuber pathogenicity ofPhytophthora infestans in the Netherlands, Potato Research, vol.30, issue.1, pp.345-354, 1998.
DOI : 10.1007/BF02358967

W. Flier, G. Van-den, L. Bosch, and . Turkensteen, Stability of partial resistance in potato cultivars exposed to aggressive strains of Phytophthora infestans, Plant Pathology, vol.7, issue.3, pp.326-337, 2003.
DOI : 10.1046/j.1365-3059.2000.00515.x

H. Flor, Current Status of the Gene-For-Gene Concept, Annual Review of Phytopathology, vol.9, issue.1, pp.275-296, 1971.
DOI : 10.1146/annurev.py.09.090171.001423

H. Pellé, G. Platt, T. Mackenzie, L. Tarn, D. Colon et al., Stability of resistance to Phytopthorra infestans in potato: an internacional evaluation, pp.364-372, 2005.

W. Fry, Phytophthora infestans: the plant (and R gene) destroyer, Molecular Plant Pathology, vol.97, issue.3, pp.385-402, 2008.
DOI : 10.1111/j.1365-3059.2007.01618.x

W. Fry and S. Goodwin, Re-emergence of Potato and Tomato Late Blight in the United States, Plant Disease, vol.81, issue.12, pp.1349-1357, 1997.
DOI : 10.1094/PDIS.1997.81.12.1349

J. Gabriel, F. Forqueda, G. Plat, and E. Fernández-northcote, Caracterización de genotipos de papa de Europa y Latinoamérica por resistencia a tizón y propiedades culinarias, Revista latinoamericana de la Papa, vol.14, issue.1, pp.10-23, 2007.

A. Gabriel, G. Coca, J. Plata, and J. Parlevliet, Characterization of the resistance to Phytophthora infestans in local potato cultivars in Bolivia, Euphytica, vol.2, issue.Suppl, pp.321-328, 2007.
DOI : 10.1007/s10681-006-9237-x

S. Goodwin, L. Sujkowski, and W. Fry, Rapid Evolution of Pathogenicity Within Clonal Lineages of the Potato Late Blight Disease Fungus, Phytopathology, vol.85, issue.6, pp.669-676, 1995.
DOI : 10.1094/Phyto-85-669

J. Henfling and W. , Field screening procedures to evaluate resistence to late blight. CIP Technology Evaluation Series No. 1982-5, 1982.

W. Henfling, El tizón tardío de la papa. 415 p. CIP Centro internacional de la papa. Boletines de información técnica. Capacitaciones y comunicaciones CIP, 1987.

R. Hijmans, G. Forbes, and T. Walter, Estimating the global severity of potato late blight with GIS-linked disease forecast models, Plant Pathology, vol.23, issue.6, pp.697-705, 2000.
DOI : 10.1175/1520-0442(1999)012<0829:RTCSTC>2.0.CO;2

M. Huarte and S. Capezio, Niveles disponibles de resistencia al tizón tardío. idiaXXI. Revista de información sobre investigación y desarrollo agropecuario, pp.101-107, 2003.

S. Jaramillo, Monografía sobre tizón tardío, 2003.

J. C. Jenkins and R. K. Jones, in Minnesota, Plant Disease, vol.87, issue.8, pp.983-990, 2003.
DOI : 10.1094/PDIS.2003.87.8.983

J. Kalazich, Variedades de papa, Manual de producción de papa para la agricultura familiar campesina (AFC). Boletín N°147, pp.49-64, 2006.

H. Lozoya-saldaña, L. Guzman-galindo, S. Fernández-pavia, E. Grünwald, and M. , Phytophthora infestans (Mont.) de Bary. I. Host-pathogen specificity and resistance components, Agrociencia, vol.40, issue.2, pp.205-217, 2006.

F. Lucca, C. Bedogni, P. Cicori, M. Huarte, and S. Capezio, Comportamiento de genotipos de Solanum tarijense, S. Gourlayi y S. tuberosum ssp. andigena frente al tizón tardío en el se bonaerense argentino, En: III Congreso Iberoamericano de Investigación y Desarrollo en Patata, pp.49-51, 2008.

H. Mendoza, Aspectos de la evaluación de la resistencia a la rancha, Phytophthora infestans. Memorias del XXIV Congreso Latinoamericano de la Papa. Asociación Latinoamericana de la papa ALAP y I Simposium Internacional de Recursos genéticos de la Papa, Instituto Nacional de Innovación Agraria -INIA, 2010.

S. Micheletto, R. Boland, and M. Huarte, Argentinian wild diploid Solanum species as sources of quantitative late blight resistance, TAG Theoretical and Applied Genetics, vol.101, issue.5-6, pp.902-906, 2000.
DOI : 10.1007/s001220051560

S. Micheletto, M. Andreoni, and M. Huarte, Vertical resistance to late blight in wild potato species from Argentina, Euphytica, vol.110, issue.2, pp.133-138, 1999.
DOI : 10.1023/A:1003793227338

O. Navia, . R. Naerstad, A. Hermansen, and T. Bjor, Exploiting host resistance to reduce the use of fungicides to control potato lete bight, Plant Pathology, vol.56, pp.156-166, 2001.

R. Naerstad, A. Hermansen, and T. Bjor, Exploiting host resistance to reduce the use of fungicides to control potato late blight, Plant Pathology, vol.78, issue.1, pp.156-166, 2007.
DOI : 10.1111/j.1365-3059.2006.01491.x

L. Ríos, El tizón tardío. Una amenaza permanente. Revista Asociación Chilena de la Papa N° 3 p, pp.13-14, 1999.

J. E. Parlevliet, Durability of resistance against fungal, bacterial and viral pathogens; present situation, Euphytica, vol.124, issue.2, pp.147-156, 2002.
DOI : 10.1023/A:1015601731446

O. Páez, R. Valverde, L. Gómez, and A. Brenes, Diversidad genética de aislamientos de Phytophthora infestans en plantaciones de papa en Costa Rica con el uso de RAPDS, Agronomía Costarricense, vol.29, issue.1, pp.41-55, 2005.

W. Pérez and G. Forbes, El tizón tardío de la papa (manual técnico) Disponible en http, 2008.

W. Pérez, S. Gamboa, M. Falcon, R. Coca, R. Raymundo et al., Genetic structure of Peruvian populations of Phytophthora infestans, Phytopathol, issue.10, pp.91-956, 2001.

J. B. Ristaino, Historical and Current Population Biology of late blight pathogen: from the Andes to the Irish Potato famine. En XV Congreso Latinoamericano de Fitopatología. Asociación Latinoamericana de Fitopatología, pp.12-16, 2009.

R. Robinson, Return to resistance: Breeding crops to reduce pesticide dependence, David California (USA) Ag Access, 1996.
DOI : 10.1016/S0167-8809(97)87015-6

G. Secor, V. Rivera, A. Thomson, and J. Rodriguez, The continuing battle for late blight resistance potato cultivars, 2009.

C. Santiago, 12-16 de enero de, 2009.

. Rodriguez, The continuing battle for late blight resistance potato cultivars. En XV Congreso Latinoamericano de Fitopatología, pp.12-16, 2009.

I. Simko, S. Costanzo, V. Ramanjulu, B. Christ, and K. Haynes, Mapping polygenes for tuber resistance to late blight in a diploid Solanum phureja ?? S. stenotomum hybrid population, Plant Breeding, vol.79, issue.4, pp.385-389, 2006.
DOI : 10.1007/s10681-005-3444-8

S. Inc, SPSS Base 15.0 user´s guide, 2006.

B. J. Staskawiez, F. M. Ausubel, B. J. Baker, J. G. Ellis, and J. D. Jones, Molecular genetics of plant disease resistance, Science, vol.268, issue.5211, pp.661-667, 1995.
DOI : 10.1126/science.7732374

K. Swiezynski, L. Domanski, H. Zarzycka, and E. Zimnoch-guzowska, The reaction of potato differentials to Phytophthora infestans isolates collected in nature, Plant Breeding, vol.9, issue.1, pp.119-126, 2000.
DOI : 10.1046/j.1439-0523.2000.00461.x

H. Torres, Manual de las enfermedades más importantes de la papa en el Perú. Centro Internacional de la papa, 2002.

G. Trujillo, Desarrollo de marcadores SCAR y CAPS en un QTL con efecto importante sobre la resistencia al tizón tardío de la papa, 2004.

L. J. Turkensteen and W. G. Flier, Late blight: Its global status in, 2002.

V. Umaerus and M. Umaerus, Inheritance of resistance to late blight, Potato Genetics. CAB Internacional, pp.365-401, 1994.

M. Visker, Association between late blight resístanse and foliage maturity type in potato. Physiological and genetic studies, 2005.

M. Visker, L. Keizer, D. Budding, L. Van-loon, L. Colon et al., Leaf Position Prevails Over Plant Age and Leaf Age in Reflecting Resistance to Late Blight in Potato, Phytopathology, vol.93, issue.6, pp.666-674, 2003.
DOI : 10.1094/PHYTO.2003.93.6.666

R. L. Wastie, Breeding for reistancePhytophthora infestans, the cause of late blight of potato, Advances in Plant Pathology, vol.7, pp.193-224, 1991.

J. E. Yuen and G. A. Forbes, in Potato Genotypes, Phytopathology, vol.99, issue.6, pp.782-786, 2009.
DOI : 10.1094/PHYTO-99-6-0782

D. Zlesak and C. Thill, Foliar resistance toPhytophthora infestans (Mont.) de Bary (US-8) in 13 Mexican and South AmericanSolarium species having EBNs of 1, 2, and 4 and implications for breeding, American Journal of Potato Research, vol.79, issue.6, pp.421-429, 2004.
DOI : 10.1007/BF02870203

L. Zúñiga, J. Molina, M. Cadena, and A. Rivera, Resistencia al tizón tardío de la papa (Phytophthora infestans) en cruzamientos de cultivares y clones de papa (Solanum tuberosum L.). Revista Mexicana de Fitopatología, pp.1-9, 2000.

H. and R. J. Spooner, Geographic distribution of wild potato species, 2001.

H. , R. J. , M. Macobs, J. Bamberg-&-d, and . Spooner, Frost tolerance in wild potato species: Assessing the predictivity of taxonomic, geographic and ecological factors, Euphytica, vol.130, pp.47-59, 2003.

J. and S. A. Hanneman, Potential gene flow between cultivated potato and its wild tuber-bearing relatives: implications for risk assessment of transgenic potatoes, 1996.

M. , C. , T. Stuessy-&-c, and . Baeza, Catálogo de la flora vascular de la, Gayana Botánica, vol.55, issue.2, pp.187-211, 1998.

R. , S. , and T. R. Stuessy-&-r, Chromosome numbers from the flora of the Juan Fernández Islands, American Journal of Botany, vol.70, issue.6, pp.799-810, 1983.

S. , T. F. , U. Swenson, D. J. Crawford, and G. S. Anderson-&-m, Plant conservation in the Juan Fernandez Archipelago, Chile, Aliso, vol.16, pp.89-98, 1998.

V. , J. P. , G. Brigneti, F. Salazar, E. W. Pehue-&-r et al., Interaction of the Solanum spp. of the Etuberosa group and nine potato-infecting viruses and viroid, Annual Applied Biology, vol.120, pp.301-313, 1992.

. Figura, Invernadero de policarbonato: producción de plantas adultas en maceteros, b) producción de flores, c) frutos en invernadero (Fotos

J. Z. Huaman, W. Williams, N. Salhuana, and . Vincent, Descriptors for the cultivated potato and for the maintenance and distribution of germoplasm collections, Consultative Group on Internacional Agricultural Research. Rome. International Board for Plant Genetic Resources, p.47, 1977.

A. Name, Meca gato blanco TUBER Shape: elongate Unusual shape: absent Predominant skin colour: white-cream Secondary skin colour: absent Distribution of secondary skin colour: absent Tuber skin type: smooth Predominant flesh colour: cream Secondary flesh colour: absent Distribution of secondary flesh colour: absent Note of eyes per tuber: intermediate Average number of eyes